Nabozhenko, Maxim, and Kiyoshi Ando. 2018. “Subtribal, Generic and Subgeneric Composition of Darkling Beetles of the Tribe Helopini (Coleoptera: Tenebrionidae) in the Eastern Palaearctic”. Acta Zoologica Academiae Scientiarum Hungaricae 64 (4), 277-327. https://doi.org/10.17109/AZH.64.4.277.2018.